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{ <http://www.tkuhn.ch/bel2nanopub/RAYfkt9uNkIM-u22DVrxIlDXul8g8HWQ_W0o59qdUR8_0#_6> ?p ?o ?g. }
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The activated TGF-beta R1 phosphorylates SMAD2 and SMAD3, which bind to the SMAD4 mediator to move into the nucleus and form complexes that regulate transcription.
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